This equation governs the dynamics of a single unknown as it evolves. In practice, there are often multiple unknowns that interact over time and one wishes to model the dynamics as the system evolves.
22 A: Lotka-Volterra equations
For example, one may consider the so-called Lotka–Volterra predator–prey model describing the evolution of a biological system involving two species: a prey population of size x(t) at time t, and a predator population of size y(t) at time t. These populations are modelled by the following system of equations: x(0) = x_0, y(0) = y_0 for some initial data (x_0,y_0) and
\begin{align}
x'(t) &= A x(t) - B x(t) y(t) \nonumber\\
y'(t) &= -C y(t) + D x(t) y(t) \tag{LV}
\end{align}
for some constants A, B, C, D \geq 0
A describes the growth rate of the prey population (this growth is proportional to the current size of the population),
B describes the rate at which the predators eat the prey (which is larger if there are more predators),
C is the death rate of the predators,
D is the rate at which predators increase due to availability of prey.
Exercise 1. Write (\text{LV}) as a problem of the form: seek u : [0,T] \to \mathbb R^2 such that
for some f: [0,T] \times \mathbb R \to \mathbb R^2.
Answer.
On defining u_1(t) := x(t) and u_2(t) := y(t), we find that
\begin{align}
u(0) &= \begin{pmatrix}x_0\\y_0\end{pmatrix} \nonumber\\
u'(t) &= \begin{pmatrix}
A u_1(t) - B u_1(t) u_2(t) \\
-C u_2(t) + D u_1(t) u_2(t)
\end{pmatrix}.
\end{align}
Notice that you may apply the numerical schemes from lectures to this system of IVPs. For example, the midpoint method reads:
functionmidpoint( u0, f, T, n ) h = T/n t =0:h:T u =fill(float(u0), n+1) u[1] = u0for j =1:n u[j+1] = u[j] + h *f( t[j] + h/2, u[j] + (h/2)*f(t[j], u[j]) )endreturn uend
midpoint (generic function with 1 method)
Here, is the numerical solution of (\text{VP}) with a particular choice of parameters A, B, C, D, initial condition u_0, and mesh \{ t_j \}:
A, B, C, D =1., 1., .5, 1.u0 = [1.,.01]f(t, u) = [A*u[1]-B*u[1]*u[2], -C*u[2]+D*u[1]*u[2]]T, n =50., 500u =midpoint( u0, f, T, n)u = [U[j] for U ∈ u, j ∈1:2]plot(0:T/n:T, u, label=["prey""predator"], l=(1, :black), m=2)
Exercise 2. Explain this plot with reference to the physical system that is being modelled.
Answer.
Initially, there are very few predators so the prey are free to grow. Eventually, this means there is available food for the predators and so the predator population grows as a result. Then, the large size of the predator population needs a large amount of food and so the prey is eaten and the prey population decreases rapidly. This reduces the available food for the predators and so the predator population also drops. This behaviour seems to converge to a stable periodic solution.
Another way to see the dynamics is to plot (x(t), y(t)) in phase space:
X, Y = [], []anim =@animatefor t ∈1:n+1 x,y = u[t,:]plot(X, Y, title="LV numerical solution", xlabel=L"x(t)", ylabel=L"y(t)", legend=false)push!(X, x)push!(Y, y)scatter!([x], [y], m=(5, :red))endmp4(anim, "pics/LV.gif")
[ Info: Saved animation to c:\Users\math5\Math 5485\Math5486\pics\LV.gif
Exercise 3. Play around with the parameters in this model (A, B, C, D). What do you notice about the numerical solution?
Answer.
There are many possible answers here - you should have played around with the following parameters and noticed that the effect makes sense when compared to the physical meaning of the variables:
A describes the growth rate of the prey population (this growth is proportional to the current size of the population),
B describes the rate at which the predators eat the prey (which is larger if there are more predators),
C is the death rate of the predators,
D is the rate at which predators increase due to availability of prey.
Exercise 4. What happens when (x_0, y_0) = (C/D, A/B)? Why?
Answer.
In this case, we have u'(t) = 0 and so u(t) = \begin{pmatrix} C/D \\ A/B \end{pmatrix} is a constant solution.
22.1 B. Higher order IVPs
Consider the follwing initial value problem: seek \theta: [0,1] \to \mathbb R such that
This is a second order IVP modelling the motion of a pendulum of length L and with damping \gamma > 0. Here, \theta is the angle made with the vertical position and g \approx 9.81 m/s^2 is the gravitational constant. See the beginning of Chapter 5 of Burden, Faires, and Burden (2015) for a picture.
Exercise 5. Show that \theta(t) = 0 is a solution when \theta_0 = 0. Explain why this makes sense with reference to the physical system we are modelling.
Answer.
If \theta(t) = 0, we have \theta''(t) = \theta'(\theta) = 0 and so - \gamma \theta'(t) -\frac{g}{L} \sin \theta(t) = 0. When the initial position of the pendulum is vertical, the pendulum will stay at rest.
Exercise 6. Write down the system of differential equations governing the quantity